1971 | B.S. | Dept. Zool., Natl. Taiwan Univ. |
1975 | Ph.D. | Dept. Biol., Univ. Rochester, USA |
1975-78 | Dept. Biol., Yale Univ., USA | |
1978-86 | Assist. & Assoc. Prof., Washinton Univ. St. Louis, USA | |
1986-2007 | Member, Div. Basic Sci., Fred Hutchinson Cancer Res. Ctr., USA | |
2004-2013 | Director & Distinguished Research Fellow, IMB | |
2013-2019 | Distinguished Research Fellow, IMB | |
2019-present | Distinguished Visiting Chair, IMB |
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